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Landscape classes are “functional units” and within each unit N inputs, plant N uptake, soils and flow pathways should be similar, though for convenience classes are often based on land use and/or soil type. All land-based processes are calculated for a generic 1 km2 cell for each landscape class within each sub-catchment. Water, nitrate and ammonium outputs from the 1 km2 cell for each land class are multiplied by the land class area, and summed to provide total inputs from the sub-catchment to the reach. These inputs are assumed to enter the stream reach directly, rather than being routed spatially from one land class to another. 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