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The QUESTOR model has sets of equations to simulates nitrogen and phosphorus species, pH and suspended sediment, the aspects of water quality of particular interest are those determinands indicative of the impacts on the river ecosystem, namely temperature, chlorophyll-a (Chl-a), BOD and DO. Chl-a is used as a surrogate for the algae using a fixed stoichiometry model whereby the ratio chl-a:C:N:P is 1:50:10:1. The growth of algae is limited by light, temperature and nutrient concentrations.  The model was developed to help catchment managers assess the impact of their actions e.g. changing discharges consents, abstracting more water from a river or building a flood relief channel. 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Landscape classes are “functional units” and within each unit N inputs, plant N uptake, soils and flow pathways should be similar, though for convenience classes are often based on land use and/or soil type. All land-based processes are calculated for a generic 1 km2 cell for each landscape class within each sub-catchment. Water, nitrate and ammonium outputs from the 1 km2 cell for each land class are multiplied by the land class area, and summed to provide total inputs from the sub-catchment to the reach. These inputs are assumed to enter the stream reach directly, rather than being routed spatially from one land class to another. 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