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Great tits and blue tits were additionally fitted with a plastic leg ring containing a RFID tag (IB Technologies, UK). All nestlings were also fitted with these tags at 2 weeks old. Additional ringing was carried out throughout the autumn and winter across the woodland. Birds were trapped using mistnets; biometrics (mass, wing length, tarsus length, fats score) were recorded for all birds and immigrants to the population were fitted with tags. All birds were released after they were processed. 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