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Data were collected from 25th April-12th May and 21st May-15th June 2013 at four sites on the Isle of May, Scotland. A grid was superimposed onto a photograph of each site. Grid cells were then classified as breeding or pre-breeding areas according to the presence or absence of breeding activity at any point during data collection (i.e. an egg or chick). A total of 69 randomly selected and individually-marked birds were followed using a telescope for 10 minute periods and their location in these grids was recorded every 15 seconds. This work was part of a NERC-funded PhD project looking at interactions between avian colonial social structure and tick-borne pathogen dynamics. 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Grid cells with ledges and no apparent evidence of breeding activity but consistent breeding activity completely surrounding them were also assumed to be breeding areas. Individual birds were identified using unique colour rings which were visible at a distance through a telescope with a x20-60 zoom lens. 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